Staff Profile
Photo for Pat Neale


Research Interests

I study the effects of sunlight, especially ultraviolet radiation, on ecosystems and the environment.  I mainly work in freshwater and marine systems focusing on the responses of the smallest organisms in those systems, the plankton.  A long term project has been studying the effects of increased UV-B due to Antarctic ozone depletion on the productivity of the Southern Ocean which surrounds the Antarctic continent.  I also supervise the SI solar-radiation monitoring program which has been measuring UV-B radiation in Maryland since the 1970s compiling one of the longest records of spectral irradiance in the world. 


  • Neale, Patrick J. ; Heaney, S. Ivan ; Jaworski, George H. M. (1991). Response to high irradiance contribute to the decline of the spring diatom maximum . Limnology and Oceanography, 36 (4) , 761-768.
  • Neale, Patrick J. ; Melis, Anastasios. (1991). Dynamics of Photosystem II heterogeneity during photoinhibition: Depletion of PS IIb from non-appressed thylakoids during strong-irradiance exposure of Chlamydomonas reinhardtii. Biochimica et biophysica acta, 1056 , 195-203.
  • Neale, Patrick J. ; Talling, Jack F. ; Heaney, S. Ivan ; Reynolds, Colin S. ; Lund John, W. G. (1991). Long time series from the English Lake District: Irradiance-dependent phytoplankton dynamics during the spring maximum . Limnology and Oceanography, 36 (4) , 751-760.
  • Neale, Patrick J. ; Melis, Anastasios. (1990). Activation of a reserve pool of photosystem II in Chlamydomonas reinhardtii counteracts photoinhibition . Plant Physiology, 92 , 1196-1204.
  • Neale, Patrick J. ; Cullen, John J. ; Yentsch, Clarice M. (1989). Bio-optical inferences from chlorophyll a fluorescence: What kind of fluorescence is measured in flow cytometry? . Limnology and Oceanography, 34 (8) , 1739-1748.
  • Demeter, Sandor ; Neale, Patrick J. ; Melis, Anastasios. (1987). Photoinhibition: Impairment of the primary charge separation between P-680 and pheophytin in photosystem II of chloroplasts. Federation of European Biochemical Societies Letters, 214 (2) , 370-374.
  • Neale, Patrick J. (1987). Algal photoinhibition and photosynthesis in the aquatic environment. Photoinhibition (39-65) Elsevier .
  • Neale, Patrick J. ; Melis, Anastasios. (1987). Response of Chlamydomonas reinhardtii reaction center stoichiometry to light intensivy variation: A spectroscopic and LDS-page analysis. Progress in Photosynthesis Research (11-11) Martinus Nijhoff Publishers .
  • Neale, Patrick J. ; Richerson, Peter J. (1987). Photoinhibition and the diurnal variation of phytoplankton photosynthesis - I. Development of a photosynthesis - irradiance model from studies of in situ responses. Journal of Plankton Research, 9 (1) , 167-193.
  • Neale, Patrick J. ; Melis, Anastasios. (1986). Algal photosynthetic membrane complexes and the photosynthesis-irradiance curve: A comparison of light-adaptation responses in Chlamydomonas reinhardtii (Chlorophyta). Journal of Phycology, 22 , 531-538.
  • Richerson, Peter J. ; Neale, Patrick J. ; Wurtsbaugh, Wayne ; Vincent, Rene Alfaro ; Vincent, Warwick. (1986). Patterns of temporal variation in Lake Titicaca. A high altitude tropical lake. I. Background, physical and chemical processes, and primary production. Hydrobiologia, 138 , 205-220.
  • Neale, Patrick J. ; Marra, John. (1985). Short-term variation of Pmax under natural irradiance conditions: A model and its implications. Marine Ecology Progress Series, 264 , 113-124.
  • Vincent, Warwick F. ; Neale, Patrick J. ; Richerson, Peter J. (1984). Photoinhibition: Algal response to bright light during diel stratification and mixing in a tropical alpine lake . Journal of Phycology, 20 , 201-211.