Staff Profile
Photo for Pat Neale


Research Interests

I study the effects of sunlight, especially ultraviolet radiation, on ecosystems and the environment.  I mainly work in freshwater and marine systems focusing on the responses of the smallest organisms in those systems, the plankton.  A long term project has been studying the effects of increased UV-B due to Antarctic ozone depletion on the productivity of the Southern Ocean which surrounds the Antarctic continent.  I also supervise the SI solar-radiation monitoring program which has been measuring UV-B radiation in Maryland since the 1970s compiling one of the longest records of spectral irradiance in the world. 


  • Williamson, Craig E., Overholt, Erin P., Brentrup, Jennifer A., Pilla, Rachel M., Leach, Taylor H., Schladow, S. G., Warren, Joseph D., Urmy, Samuel S., Sadro, Steven, Chandra, Sudeep and Neale, Patrick J. (2016). Sentinel responses to droughts, wildfires, and floods: effects of UV radiation on lakes and their ecosystem services. Frontiers in Ecology and the Environment, 14 (2) , 102-109.
  • Gallegos, Charles L. and Neale, Patrick J. (2015). Long-term variations in primary production in a eutrophic sub-estuary: Contribution of short-term events. Estuarine, Coastal and Shelf Science, 162 , 22-34.
  • Sanders, Robert W., Cooke, Sandra L., Fischer, Janet M., Fey, Samuel B., Heinze, Adam W., Jeffrey, Wade H., Macaluso, Amy L., Moeller, Robert E., Morris, Donald P., Neale, Patrick J., Olson, Mark H., Dean Pakulski, J., Porter, Jason A., Schoener, Donald M. and Williamson, Craig E. (2015). Shifts in microbial food web structure and productivity after additions of naturally occurring dissolved organic matter: Results from large-scale lacustrine mesocosms. Limnology and Oceanography, 60 (6) , 2130-2144.
  • Thomas, Brian C., Neale, Patrick J. and Snyder, Brock R. (2015). Solar Irradiance Changes and Photobiological Effects at Earth's Surface Following Astrophysical Ionizing Radiation Events. Astrobiology, 15 (3) , 207-220.
  • Figueroa, F. L., Mercado, J. M., Beardall, J., Neale, Patrick J., Montecino, V. and Kromkamp, J. C. (2014). Introduction. Aquatic Biology, 22 , 1-4.
  • Fragoso, Glaucia Moreira, Neale, Patrick J., Kana, Todd Milan and Pritchard, Alicia Lynn. (2014). Kinetics of Photosynthetic Response to Ultraviolet and Photosynthetically Active Radiation in Synechococcus WH8102 (CYANOBACTERIA). Photochemistry and photobiology, 90 (3) , 522-532.
  • Kieber, David J., Miller, Gary W., Neale, Patrick J. and Mopper, Kenneth. (2014). Wavelength and temperature-dependent apparent quantum yields for photochemical formation of hydrogen peroxide in seawater. Environmental Science: Processes & Impacts, 16 (4)
  • Mercado, J. M., Sobrino, C., Neale, Patrick J., Segovia, M., Reul, A., Amorim, A. L., Carrillo, P., Claquin, P., Cabrerizo, M. J., Leon, P., Lorenzo, M. R., Medina-Sanchez, J. M., Montecino, V., Napoleon, C., Prasil, O., Putzeys, S., Salles, S. and Yebra, L. (2014). Effect of CO2, nutrients and light on coastal plankton. II. Metabolic rates. Aquatic Biology, 22 , 43-57.
  • Neale, Patrick J., Sobrino, C., Segovia, M., Mercado, J. M., Leon, P., Cortes, M. D., Tuite, P., Picazo, A., Salles, S., Cabrerizo, M. J., Prasil, O., Montecino, V., Reul, A. and Fuentes-Lema, A. (2014). Effect of CO2, nutrients and light on coastal plankton. I. Abiotic conditions and biological responses. Aquatic Biology, 22 , 25-41.
  • Reul, A., Munoz, M., Bautista, B., Neale, Patrick J., Sobrino, C., Mercado, J. M., Segovia, M., Salles, S., Kulk, G., Leon, P., van de Poll, W. H., Perez, E., Buma, A. and Blanco, J. M. (2014). Effect of CO2, nutrients and light on coastal plankton. III. Trophic cascade, size structure and composition. Aquatic Biology, 22 , 59-76.
  • Sobrino, C., Segovia, M., Neale, Patrick J., Mercado, J. M., Garcia-Gomez, C., Kulk, G., Lorenzo, M. R., Camarena, T., van de Poll, W. H., Spilling, K. and Ruan, Z. (2014). Effect of CO2, nutrients and light on coastal plankton. IV. Physiological responses. Aquatic Biology, 22 , 77-93.
  • Neale, Patrick J. (2013). Species Specific Responses to Combined Thermal-irradiance Stress in Microalgae - Each is to its Own. Photochemistry and Photobiology, 89 (4) , 822-823.
  • Neale, Patrick J., Pritchard, A. L. and Ihnacik, R. (2013). UV effects on the primary productivity of picophytoplankton: biological weighting functions and exposure response curves of Synechococcus. Biogeosciences Discussions, 10 (12) , 19449-19484.
  • Neale, Patrick J., Sobrino, Cristina and Gargett, Ann E. (2012). Vertical Mixing and the Effects of Solar Radiation on Photosystem II Electron Transport by Phytoplankton in the Ross Sea Polynya. Deep Sea Research Part I: Oceanographic Research Papers, 63 , 118-132.
  • Smyth, Robyn L., Sobrino, Cristina, Phillips-Kress, Jesse, Kim, Hae-Cheol and Neale, Patrick J. (2012). Phytoplankton photosynthetic response to solar ultraviolet irradiance in the Ross Sea Polynya: Development and evaluation of a time-dependent model with limited repair. Limnology and Oceanography, 57 (6) , 1602-1618.