Staff Profile
Photo for Pat Neale


Research Interests

I study the effects of sunlight, especially ultraviolet radiation, on ecosystems and the environment.  I mainly work in freshwater and marine systems focusing on the responses of the smallest organisms in those systems, the plankton.  A long term project has been studying the effects of increased UV-B due to Antarctic ozone depletion on the productivity of the Southern Ocean which surrounds the Antarctic continent.  I also supervise the SI solar-radiation monitoring program which has been measuring UV-B radiation in Maryland since the 1970s compiling one of the longest records of spectral irradiance in the world. 


  • Neale, Patrick J. and Thomas, Brian C. (2017). Inhibition by Ultraviolet and Photosynthetically Available Radiation Lowers Model Estimates of Depth-Integrated Picophytoplankton Photosynthesis: Global Predictions for Prochlorococcus and Synechococcus. Global Change Biology, 23 (1) , 293-306.
  • Nelson, Natalie G., Muñoz-Carpena, Rafael, Neale, Patrick J., Tzortziou, Maria and Megonigal, J. Patrick. (2017). Temporal variability in the importance of hydrologic, biotic, and climatic descriptors of dissolved oxygen dynamics in a shallow tidal-marsh creek. Water Resources Research, 53 (8) , 7103-7120.
  • Smyth, Robyn L., Akan, Cigdem, Tejada-Martínez, Andrés and Neale, Patrick J. (2017). Quantifying phytoplankton productivity and photoinhibition in the Ross Sea Polynya with large eddy simulation of Langmuir circulation. Journal of Geophysical Research: Oceans, 122 (7) , 5545-5565.
  • Whigham, Dennis F., Walker, C. M., Maurer, J., King, R. S., Hauser, W., Baird, S., Keuskamp, J. A. and Neale, Patrick J. (2017). Watershed influences on the structure and function of riparian wetlands associated with headwater streams – Kenai Peninsula, Alaska . Science of The Total Environment, 599–600 , 124-134.
  • Neale, Patrick J. and Thomas, Brian C. (2016). Solar Irradiance Changes and Phytoplankton Productivity in Earth's Ocean Following Astrophysical Ionizing Radiation Events. Astrobiology, 16 (4) , 245-258.
  • Williamson, Craig E., Overholt, Erin P., Brentrup, Jennifer A., Pilla, Rachel M., Leach, Taylor H., Schladow, S. G., Warren, Joseph D., Urmy, Samuel S., Sadro, Steven, Chandra, Sudeep and Neale, Patrick J. (2016). Sentinel responses to droughts, wildfires, and floods: effects of UV radiation on lakes and their ecosystem services. Frontiers in Ecology and the Environment, 14 (2) , 102-109.
  • Gallegos, Charles L. and Neale, Patrick J. (2015). Long-term variations in primary production in a eutrophic sub-estuary: Contribution of short-term events. Estuarine, Coastal and Shelf Science, 162 , 22-34.
  • Sanders, Robert W., Cooke, Sandra L., Fischer, Janet M., Fey, Samuel B., Heinze, Adam W., Jeffrey, Wade H., Macaluso, Amy L., Moeller, Robert E., Morris, Donald P., Neale, Patrick J., Olson, Mark H., Dean Pakulski, J., Porter, Jason A., Schoener, Donald M. and Williamson, Craig E. (2015). Shifts in microbial food web structure and productivity after additions of naturally occurring dissolved organic matter: Results from large-scale lacustrine mesocosms. Limnology and Oceanography, 60 (6) , 2130-2144.
  • Thomas, Brian C., Neale, Patrick J. and Snyder, Brock R. (2015). Solar Irradiance Changes and Photobiological Effects at Earth's Surface Following Astrophysical Ionizing Radiation Events. Astrobiology, 15 (3) , 207-220.
  • Figueroa, F. L., Mercado, J. M., Beardall, J., Neale, Patrick J., Montecino, V. and Kromkamp, J. C. (2014). Introduction. Aquatic Biology, 22 , 1-4.
  • Fragoso, Glaucia Moreira, Neale, Patrick J., Kana, Todd Milan and Pritchard, Alicia Lynn. (2014). Kinetics of Photosynthetic Response to Ultraviolet and Photosynthetically Active Radiation in Synechococcus WH8102 (CYANOBACTERIA). Photochemistry and photobiology, 90 (3) , 522-532.
  • Kieber, David J., Miller, Gary W., Neale, Patrick J. and Mopper, Kenneth. (2014). Wavelength and temperature-dependent apparent quantum yields for photochemical formation of hydrogen peroxide in seawater. Environmental Science: Processes & Impacts, 16 (4)
  • Mercado, J. M., Sobrino, C., Neale, Patrick J., Segovia, M., Reul, A., Amorim, A. L., Carrillo, P., Claquin, P., Cabrerizo, M. J., Leon, P., Lorenzo, M. R., Medina-Sanchez, J. M., Montecino, V., Napoleon, C., Prasil, O., Putzeys, S., Salles, S. and Yebra, L. (2014). Effect of CO2, nutrients and light on coastal plankton. II. Metabolic rates. Aquatic Biology, 22 , 43-57.
  • Neale, Patrick J., Sobrino, C., Segovia, M., Mercado, J. M., Leon, P., Cortes, M. D., Tuite, P., Picazo, A., Salles, S., Cabrerizo, M. J., Prasil, O., Montecino, V., Reul, A. and Fuentes-Lema, A. (2014). Effect of CO2, nutrients and light on coastal plankton. I. Abiotic conditions and biological responses. Aquatic Biology, 22 , 25-41.
  • Reul, A., Munoz, M., Bautista, B., Neale, Patrick J., Sobrino, C., Mercado, J. M., Segovia, M., Salles, S., Kulk, G., Leon, P., van de Poll, W. H., Perez, E., Buma, A. and Blanco, J. M. (2014). Effect of CO2, nutrients and light on coastal plankton. III. Trophic cascade, size structure and composition. Aquatic Biology, 22 , 59-76.