A decadal study of headwater streams has involved SERC and partner organizations (Baylor University, University of South Florida, Kachemak Bay Research Reserve) with funding from EPA and the Alaska Salmon Fund. The project has focused on headwater streams that are too small for adult salmon to reach but serve as rearing habitat for millions of juvenile salmon that spend from 1-3 years in the streams before migrating to the sea. The project has demonstrated that the streams fall into two broad functional groups (high and low gradient) based primarily on the slope of the stream. The study has also demonstrated strong linkages between the land and the streams with more than 60% of the nitrogen in the bodies of juvenile salmon coming from the land. Primary and secondary production of the streams appears to be either nitrogen or carbon limited which is a function of whether or not alder (a nitrogen fixing plant) is present on the watershed and how much of the watershed consists of large peat-based wetlands. For the first time, it has also been demonstrated that structure and function of the streamside wetlands is strongly influenced by the presence or absence of alder on the watershed.