Staff Profile
Photo for Pat Neale


Research Interests

I study the effects of sunlight, especially ultraviolet radiation, on ecosystems and the environment.  I mainly work in freshwater and marine systems focusing on the responses of the smallest organisms in those systems, the plankton.  A long term project has been studying the effects of increased UV-B due to Antarctic ozone depletion on the productivity of the Southern Ocean which surrounds the Antarctic continent.  I also supervise the SI solar-radiation monitoring program which has been measuring UV-B radiation in Maryland since the 1970s compiling one of the longest records of spectral irradiance in the world. 


  • Neale, Patrick J., Cullen, John J. and Yentsch, Clarice M. (1989). Bio-optical inferences from chlorophyll a fluorescence: What kind of fluorescence is measured in flow cytometry? . Limnology and Oceanography, 34 (8) , 1739-1748.
  • Demeter, Sandor, Neale, Patrick J. and Melis, Anastasios. (1987). Photoinhibition: Impairment of the primary charge separation between P-680 and pheophytin in photosystem II of chloroplasts. Federation of European Biochemical Societies Letters, 214 (2) , 370-374.
  • Neale, Patrick J. (1987). Algal photoinhibition and photosynthesis in the aquatic environment. Photoinhibition (39-65) Elsevier .
  • Neale, Patrick J. and Melis, Anastasios. (1987). Response of Chlamydomonas reinhardtii reaction center stoichiometry to light intensivy variation: A spectroscopic and LDS-page analysis. Progress in Photosynthesis Research (11-11) Martinus Nijhoff Publishers .
  • Neale, Patrick J. and Richerson, Peter J. (1987). Photoinhibition and the diurnal variation of phytoplankton photosynthesis - I. Development of a photosynthesis - irradiance model from studies of in situ responses. Journal of Plankton Research, 9 (1) , 167-193.
  • Neale, Patrick J. and Melis, Anastasios. (1986). Algal photosynthetic membrane complexes and the photosynthesis-irradiance curve: A comparison of light-adaptation responses in Chlamydomonas reinhardtii (Chlorophyta). Journal of Phycology, 22 , 531-538.
  • Richerson, Peter J., Neale, Patrick J., Wurtsbaugh, Wayne, Vincent, Rene Alfaro and Vincent, Warwick. (1986). Patterns of temporal variation in Lake Titicaca. A high altitude tropical lake. I. Background, physical and chemical processes, and primary production. Hydrobiologia, 138 , 205-220.
  • Neale, Patrick J. and Marra, John. (1985). Short-term variation of Pmax under natural irradiance conditions: A model and its implications. Marine Ecology Progress Series, 264 , 113-124.
  • Vincent, Warwick F., Neale, Patrick J. and Richerson, Peter J. (1984). Photoinhibition: Algal response to bright light during diel stratification and mixing in a tropical alpine lake . Journal of Phycology, 20 , 201-211.