Staff Profile
Photo for Pat Neale


Research Interests

I study the effects of sunlight, especially ultraviolet radiation, on ecosystems and the environment.  I mainly work in freshwater and marine systems focusing on the responses of the smallest organisms in those systems, the plankton.  A long term project has been studying the effects of increased UV-B due to Antarctic ozone depletion on the productivity of the Southern Ocean which surrounds the Antarctic continent.  I also supervise the SI solar-radiation monitoring program which has been measuring UV-B radiation in Maryland since the 1970s compiling one of the longest records of spectral irradiance in the world. 


  • Sobrino, C., Segovia, M., Neale, Patrick J., Mercado, J. M., Garcia-Gomez, C., Kulk, G., Lorenzo, M. R., Camarena, T., van de Poll, W. H., Spilling, K. and Ruan, Z. (2014). Effect of CO2, nutrients and light on coastal plankton. IV. Physiological responses. Aquatic Biology, 22 , 77-93.
  • Neale, Patrick J., Sobrino, C., Segovia, M., Mercado, J. M., Leon, P., Cortes, M. D., Tuite, P., Picazo, A., Salles, S., Cabrerizo, M. J., Prasil, O., Montecino, V., Reul, A. and Fuentes-Lema, A. (2014). Effect of CO2, nutrients and light on coastal plankton. I. Abiotic conditions and biological responses. Aquatic Biology, 22 , 25-41.
  • Kieber, David J., Miller, Gary W., Neale, Patrick J. and Mopper, Kenneth. (2014). Wavelength and temperature-dependent apparent quantum yields for photochemical formation of hydrogen peroxide in seawater. Environmental Science: Processes & Impacts, 16 (4)
  • Neale, Patrick J., Pritchard, A. L. and Ihnacik, R. (2013). UV effects on the primary productivity of picophytoplankton: biological weighting functions and exposure response curves of Synechococcus. Biogeosciences Discuss, 10 (12) , 19449-19484.
  • Neale, Patrick J. (2013). Species Specific Responses to Combined Thermal-irradiance Stress in Microalgae - Each is to its Own. Photochemistry and Photobiology, 89 (4) , 822-823.
  • Smyth, Robyn L., Sobrino, Cristina, Phillips-Kress, Jesse, Kim, Hae-Cheol and Neale, Patrick J. (2012). Phytoplankton photosynthetic response to solar ultraviolet irradiance in the Ross Sea Polynya: Development and evaluation of a time-dependent model with limited repair. Limnology and Oceanography, 57 (6) , 1602-1618.
  • Torres-Nuñez, E., Sobrino, C., Neale, Patrick J., Ceinos, R. M., Du, S. J. and Rotllant, J. (2012). Molecular Response to Ultraviolet Radiation Exposure in Fish Embryos: Implications for Survival and Morphological Development. Photochemistry and photobiology, 88 (3) , 701-707.
  • Neale, Patrick J., Sobrino, Cristina and Gargett, Ann E. (2012). Vertical Mixing and the Effects of Solar Radiation on Photosystem II Electron Transport by Phytoplankton in the Ross Sea Polynya. Deep Sea Research Part I: Oceanographic Research Papers, 63 , 118-132.
  • Tzortziou, Maria, Neale, Patrick J., Megonigal, J. Patrick, Pow, Crystal Lee and Butterworth, Megan. (2011). Spatial gradients in dissolved carbon due to tidal marsh outwelling into a Chesapeake Bay estuary . Marine Ecology Progress Series, 426 , 41-56.
  • Neale, Patrick J., Jeffrey, Wade H., Sobrino, Cristina, Pakulski, J. Dean, Phillips-Kress, Jesse, Baldwin, Amy J., Franklin, Linda and Kim, Hae-Cheol. (2009). Inhibition of Phytoplankton and Bacterial Productivity by Solar Radiation in the Ross Sea Polynya . Smithsonian at the Poles: Contributions to International Polar Year Science (299-308) .
  • Miller, Harlan L., III., Neale, Patrick J. and Dunton, Kenneth H. (2009). Biological Weighting Functions for UV inhibition of photosynthesis in the Kelp Laminaria hyperborea (Phaeophyceae). Journal of Phycology, 45 (3) , 571-584.
  • Tzortziou, Maria, Gallegos, Charles L., Neale, Patrick J., Subramaniam, Ajit, Herman, Jay R. and Harding, Lawrence W., Jr. (2009). Bio-optical characteristics and remote sensing in the mid Chesapeake Bay through intergration of observations and radiative transfer closure. Remote Sensing and Geospatial Technologies for Coastal Ecosystem Assessment and Management (139-168) Springer .
  • Moisan, T. A., Goes, J. and Neale, Patrick J. (2009). Mycosporine-Like Amino Acids In Phytoplankton: Biochemistry, Physiology, and Optics. Marine Phytoplankton (119-143) Nova Science Publishers .
  • MacKay, M. D., Neale, Patrick J., Arp, C. D., Domis, L. N. De Senerpont, Fang, X., Gal, G., Joehnk, K. D., Kirillin, G., Lenters, J. D., Litchman, E., MacIntyre, S., Marsh, P., Melack, J., Mooij, W. M., Peeters, F., Quesada, A., Schladow, S. G., Schmid, M., Spence, C. and Stokes, S. L. (2009). Modeling lakes and reservoirs in the climate system . Limnology and Oceanography, 54 (6) , 2315-2329.
  • Sobrino, Cristina, Neale, Patrick J., Phillips-Kress, J. D., Moeller, R. E. and Porter, J. A. (2009). Elevated CO2 increases sensitivity to ultraviolet radiation in lacustrine phytoplankton assemblages . Limnology and Oceanography, 54 (6) , 2448-2459.